WATCH: World leaders and veterans mark 80th anniversary of D-Day in France

OMAHA BEACH, France (AP) — As young soldiers, they waded ashore in Normandy through gunfire to battle the Nazis. On Thursday, a dwindling number of World War II veterans in a parade of wheelchairs joined a new generation of leaders to honor the dead, the living and the fight for democracy in moving commemorations on and around those same beaches where they landed exactly 80 years ago on D-Day.

Watch the ceremony live in our player above.

The war in Ukraine shadowed the ceremonies, a grim modern-day example of lives and cities that are again suffering through war in Europe.

The break of dawn eight decades after Allied troops landed on five code-named beaches — Utah, Omaha, Gold, Juno and Sword — started the day of remembrance by Allied nations now standing together again behind Ukraine.

WATCH: D-Day veterans return to Normandy for 80th anniversary of Allied invasion

World War II ally Russia, which launched a full-scale invasion of Ukraine in 2022, wasn’t invited, and U.S. President Joe Biden directly linked Ukraine’s fight for its young democracy to the battle to defeat Nazi Germany.

“To surrender to bullies, to bow down to dictators is simply unthinkable,” Biden said. “If we were to do that, it means we’d be forgetting what happened here on these hallowed beaches.”

With the dead and wounded on both sides in Ukraine estimated in the hundreds of thousands, commemorations for the more than 4,400 Allied dead on D-Day and many tens of thousands more, including French civilians, killed in the ensuing Battle of Normandy are tinged with concerns that World War II lessons are being lost.

READ MORE: D-Day anniversary shines a spotlight on ‘Rosie the Riveter’ women who built the weapons of WWII

“There are things worth fighting for,” said Walter Stitt, who fought in tanks and turns 100 in July, as he visited Omaha Beach this week. “Although I wish there was another way to do it than to try to kill each other.”

“We’ll learn one of these days, but I won’t be around for that,” he said.

As now-centenarian veterans revisited old memories and fallen comrades buried in Normandy graves, Ukrainian President Volodymyr Zelenskyy’s presence at the international D-Day commemoration was met with loud applause, fusing World War II’s awful past with the fraught present. More applause followed when a giant screen showed the Ukrainian leader speaking with U.S. veterans.

French President Emmanuel Macron awarded the Legion of Honor to 11 U.S. veterans and a British female veteran. Among the Americans was Edward Berthold, a pilot who carried out his three missions over France in May 1944, before taking part in an operation in Saint-Lo, in Normandy, on D-Day. He flew 35 combat missions in all during World War II.

“You came here because the free world needed each and every one of you, and you answered the call,” Macron said. “You came here to make France a free nation. You’re back here today at home, if I may say.”

READ MORE: Week of ceremonies for D-Day’s 80th anniversary kicks off with parachute jump over Normandy

Macron also bestowed the Legion of Honor on 103-year-old Christian Lamb, the daughter of a Royal Navy admiral who was studying in Normandy in 1939 when her father called her back to London. There, Lamb created detailed maps that guided the crews of landing craft on D-Day.

The French president bent down to Lamb in a wheelchair, pin the medal and kiss her on both cheeks, describing her as one of the “heroes in the shadows.”

Conscious of the inevitability of age and time for World War II veterans, huge throngs of aficionados in uniforms and riding vehicles of the time, and tourists soaking up the spectacle, flooded Normandy for the 80th anniversary. At the international ceremony later, the veterans received a standing ovation as they were paraded before the stands in a stately line of wheelchairs to avoid the long walk across the beachfront promenade.

“We just have to remember the sacrifices of everybody who gave us our freedom,” said Becky Kraubetz, a Briton now living in Florida whose grandfather served with the British Army during World War II and was captured in Malta. She was among a crowd of thousands of people that stretched for several kilometers (miles) along Utah Beach, the westernmost of the D-Day beaches.

In a quiet spot away from the official ceremonies, France’s Christophe Receveur performed his own tribute, unfurling an American flag he had bought on a trip to Pennsylvania to honor those who died on D-Day.

“To forget them is to let them die all over again,” the 57-year-old said as he and his daughter, Julie, then carefully refolded the flag into a tight triangle, adding that those now dying in Ukraine fighting the invading Russian army were also on his mind.

“All these troops came to liberate a country that they didn’t know for an ideology — democracy, freedom — that is under severe strain now,” he said.

For Warren Goss, a 99-year-old American veteran of D-Day who landed in the first waves on Utah Beach, a visit years later to the same place where he watched a comrade fall to incoming fire confirmed that the sacrifice was worthy.

“I looked at the beach and it was beautiful, all the people, the kids were playing and I see the boys and girls were walking, holding hands, with their life back,” he told the Danish king and prime minister, who hung on his every word.

The fair-like atmosphere fueled by World War II-era jeeps and trucks tearing down hedge-rowed lanes so deadly for Allied troops who fought dug-in German defenders, and of reenactors playing at war on sands where D-Day soldiers fell, leave open the question of what meaning anniversaries will have once the veterans are gone.

But at the 80th, they’re the VIPs of commemorations across the Normandy coast where the largest-ever land, sea and air armada punctured Hitler’s defenses in Western Europe and helped precipitate his downfall 11 months later.

A few days before D-Day, a pilot in Sergeant Melvin “Mel” Jenner’s squadron asked if Jenner would fly a top-secret mission on June 6. Accepting the invitation, Melvin was tasked with flying up and down the French coast photographing the historic invasion. 80 years later, during a visit back to Normandy, France, Jenner recounted his memories of D-Day and the heroes, he said, who gave there lives that day.

“They really were the golden generation, those 17-, 18-year-old guys doing something so brave,” said James Baker, a 56-year-old from the Netherlands, reflecting on Utah Beach.

Farther up the coast on Gold Beach, a military bagpiper played at precisely the time that British troops landed there 80 years ago.

The United Kingdom’s King Charles III and Prime Minister Rishi Sunak were among those at a ceremony to honor the troops who landed there and on Sword Beach, while Prince William and Canadian Prime Minister Justin Trudeau joined others at ceremony for the Canadian troops at Juno Beach.

In his address, the king told the crowd that the world was fortunate that a generation “did not flinch” when they were called upon.

“Our obligation to remember what they stood for and what they achieved for us all can never diminish,” he said.

Speaking in French, Charles also paid tribute to the “unimaginable number” of French civilians killed in the battle for Normandy, and the bravery and sacrifice of the French Resistance.

Those who traveled to Normandy include women who were among the millions who built bombers, tanks and other weaponry and played other vital World War II roles that were long overshadowed by the combat exploits of men.

Feted everywhere they go in wheelchairs and walking with canes, veterans are using their voices to repeat their message they hope will live eternal: Never forget.

“We weren’t doing it for honors and awards. We were doing it to save our country,” said 98-year-old Anna Mae Krier, who worked as a riveter building B-17 and B-29 bombers. “We ended up helping save the world.”

Jill Lawless in London, and Jeffrey Schaeffer, Mark Carlson, Bela Szandelszky, Helena Alves and Alex Turnbull along the Normandy coast, contributed to this report.